新托福TPO2阅读原文(一):TheOriginsofCetaceansTPO-2-1：TheOriginsofCetaceansItshouldbeobviousthatcetaceans—whales,porpoises,anddolphins—aremammals.Theybreathethroughlungs,notthroughgills,andgivebirthtoliveyoung.Theirstreamlinedbodies,theabsenceofhindlegs,andthepresenceofafluke1andblowhole2cannotdisguisetheiraffinitieswithlanddwellingmammals.However,unlikethecasesofseaottersandpinnipeds(seals,sealions,andwalruses,whoselimbsarefunctionalbothonlandandatsea),itisnoteasytoenvisionwhatthefirstwhaleslookedlike.Extinctbutalreadyfullymarinecetaceansareknownfromthefossilrecord.Howwasthegapbetweenawalkingmammalandaswimmingwhalebridged?Missinguntilrecentlywerefossilsclearlyintermediate,ortransitional,betweenlandmammalsandcetaceans.Veryexcitingdiscoverieshavefinallyallowedscientiststoreconstructthemostlikelyoriginsofcetaceans.In1979,ateamlookingforfossilsinnorthernPakistanfoundwhatprovedtobetheoldestfossilwhale.ThefossilwasofficiallynamedPakicetusinhonorofthecountrywherethediscoverywasmade.Pakicetuswasfoundembeddedinrocksformedfromriverdepositsthatwere52millionyearsold.TheriverthatformedthesedepositswasactuallynotfarfromanancientoceanknownastheTethysSea.Thefossilconsistsofacompleteskullofanarchaeocyte,anextinctgroupofancestorsofmoderncetaceans.Althoughlimitedtoaskull,thePakicetusfossilprovidespreciousdetailsontheoriginsofcetaceans.Theskulliscetacean-likebutitsjawboneslacktheenlargedspacethatisfilledwithfatoroilandusedforreceivingunderwatersoundinmodernwhales.Pakicetusprobablydetectedsoundthroughtheearopeningasinlandmammals.Theskullalsolacksablowhole,anothercetaceanadaptationfordiving.Otherfeatures,however,showexpertsthatPakicetusisatransitionalformbetweenagroupofextinctflesh-eatingmammals,themesonychids,andcetaceans.IthasbeensuggestedthatPakicetusfedonfishinshallowwaterandwasnotyetadaptedforlifeintheopenocean.Itprobablybredandgavebirthonland.AnothermajordiscoverywasmadeinEgyptin1989.Severalskeletonsofanotherearlywhale,Basilosaurus,werefoundinsedimentsleftbytheTethysSeaandnowexposedintheSaharadesert.Thiswhalelivedaround40millionyearsago,12millionyearsafterPakicetus.Manyincompleteskeletonswerefoundbuttheyincluded,forthefirsttimeinanarchaeocyte,acompletehindlegthatfeaturesafootwiththreetinytoes.Suchlegswouldhavebeenfartoosmalltohavesupportedthe50-foot-longBasilosaurusonland.Basilosauruswasundoubtedlyafullymarinewhalewithpossiblynonfunctional,orvestigial,hindlegs.Anevenmoreexcitingfindwasreportedin1994,alsofromPakistan.ThenowextinctwhaleAmbulocetusnatans("thewalkingwhalethatswam")livedintheTethysSea49millionyearsago.Itlivedaround3millionyearsafterPakicetusbut9millionbeforeBasilosaurus.Thefossilluckilyincludesagoodportionofthehindlegs.Thelegswerestrongandendedinlongfeetverymuchlikethoseofamodernpinniped.Thelegswerecertainlyfunctionalbothonlandandatsea.Thewhaleretainedatailandlackedafluke,themajormeansoflocomotioninmoderncetaceans.Thestructureofthebackboneshows,however,thatAmbulocetusswamlikemodernwhalesbymovingtherearportionofitsbodyupanddown,eventhoughaflukewasmissing.Thelargehindlegswereusedforpropulsioninwater.Onland,whereitprobablybredandgavebirth,Ambulocetusmayhavemovedaroundverymuchlikeamodernsealion.Itwasundoubtedlyawhalethatlinkedlifeonlandwithlifeatsea.译文：TPO-2-1鲸类的起源众所周知，鲸类动物是哺乳动物,如鲸鱼、鼠海豚和海豚。它们用肺呼吸，而不是鳃，属于胎生。鲸类动物呈流线型的身...
